If the filing person has previously filed a statement on Schedule 13G to report the acquisition that is the subject of this Schedule 13D, and is filing this schedule because of §§ 240.13d-1(e), 240.13d-1(f) or 240.13d-1(g), check the following box.

The information required on the remainder of this cover page shall not be deemed to be “filed” for the purpose of Section 18 of the Securities Exchange Act of 1934 (“Act”) or otherwise subject to the liabilities of that section of the Act but shall be subject to all other provisions of the Act (however, see the Notes).




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SCHEDULE 13D




Comment for Type of Reporting Person:
Consists of (i) 1,386,122 shares of Common Stock held of record by TCG III LP (as defined in Item 2(a) below), (ii) warrants to purchase 5,218 shares of Common Stock held by TCG III LP which are exercisable within 60 days of this Statement, (iii) 1,565,311 shares of Common Stock held of record by TCG III-A LP (as defined in Item 2(a) below) and (iv) warrants to purchase 5,893 shares of Common Stock held by TCG III-A LP which are exercisable within 60 days of this Statement. TCG III GP LP is the general partner of each of TCG III LP and TCG III-A LP and may be deemed to have voting, investment and dispositive power with respect to these securities. Peter Svennilson and Tim Kutzkey, a member of the Issuer's board of directors, are the managing partners of TCG III GP LP and may each be deemed to share voting, investment and dispositive power with respect to these securities. Excludes (a) warrants to purchase 594,625 shares of Common Stock held by TCG III LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations and (b) warrants to purchase 671,518 shares of Common Stock held by TCG III-A LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations. Based on 17,807,990 shares outstanding, comprised of (i) 11,753,558 shares of Common Stock outstanding as of August 4, 2026, as reported by the Issuer (as defined in Item 1(b) below) in its quarterly report on Form 10-Q filed with the Securities and Exchange Commission (the Commission) on August 6, 2026 (the Form 10-Q), plus (ii) 6,043,321 shares of Common Stock issued in the the second tranche of the Issuer's private placement which closed on October 20, 2026 (the Private Placement), as disclosed to the Reporting Persons by the Issuer, plus (iii) 5,218 shares underlying warrants to purchase shares of Common Stock held by TCG III LP which are exercisable within 60 days of this Statement, plus (iv) 5,893 shares underlying warrants to purchase shares of Common Stock held by TCG III-A LP which are exercisable within 60 days of this Statement.


SCHEDULE 13D




Comment for Type of Reporting Person:
Consists of (i) 1,386,122 shares of Common Stock held of record by TCG III LP and (ii) warrants to purchase 5,218 shares of Common Stock held by TCG III LP which are exercisable within 60 days of this Statement. TCG III GP LP is the general partner of TCG III LP and may be deemed to have voting, investment and dispositive power with respect to these securities. Peter Svennilson and Tim Kutzkey, a member of the Issuer's board of directors, are the managing partners of TCG III GP LP and may each be deemed to share voting, investment and dispositive power with respect to these securities. Excludes warrants to purchase 594,625 shares of Common Stock held by TCG III LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations. Based on 17,802,097 shares outstanding, comprised of (i) 11,753,558 shares of Common Stock outstanding as of August 4, 2026, as reported by the Issuer in the Form 10-Q, plus (ii) 6,043,321 shares of Common Stock issued in the Private Placement, as disclosed to the Reporting Persons by the Issuer, plus (iii) 5,218 shares underlying warrants to purchase shares of Common Stock held by TCG III LP which are exercisable within 60 days of this Statement.


SCHEDULE 13D




Comment for Type of Reporting Person:
Consists of (i) 1,565,311 shares of Common Stock held of record by TCG III-A LP and (ii) warrants to purchase 5,893 shares of Common Stock held by TCG III-A LP which are exercisable within 60 days of this Statement. TCG III GP LP is the general partner of TCG III-A LP and may be deemed to have voting, investment and dispositive power with respect to these securities. Peter Svennilson and Tim Kutzkey, a member of the Issuer's board of directors, are the managing partners of TCG III GP LP and may each be deemed to share voting, investment and dispositive power with respect to these securities. Excludes warrants to purchase 671,518 shares of Common Stock held by TCG III-A LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations. Based on 17,802,772 shares outstanding, comprised of (i) 11,753,558 shares of Common Stock outstanding as of August 4, 2026, as reported by the Issuer in the Form 10-Q, plus (ii) 6,043,321 shares of Common Stock issued in the Private Placement, as disclosed to the Reporting Persons by the Issuer, plus (iii) 5,893 shares underlying warrants to purchase shares of Common Stock held by TCG III-A LP which are exercisable within 60 days of this Statement.


SCHEDULE 13D




Comment for Type of Reporting Person:
Consists of 862,068 shares of Common Stock held of record by TCG Opportunity III LP (as defined in Item 2(a)). TCG Opportunity III GP LP (as defined in Item 2(a)) is the general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to these securities. TCG Opportunity III GP LLC (as defined in Item 2(a)) is the general partner of TCG Opportunity III GP LP and the ultimate general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to these securities. Peter Svennilson and Tim Kutzkey, a member of the Issuer's board of directors, are the managing members of TCG Opportunity III GP LLC and may each be deemed to share voting, investment and dispositive power with respect to these securities. Excludes warrants to purchase 431,034 shares of Common Stock held by TCG Opportunity III LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations. Based on 17,796,879 shares outstanding, comprised of (i) 11,753,558 shares of Common Stock outstanding as of August 4, 2026, as reported by the Issuer in the Form 10-Q, plus (ii) 6,043,321 shares of Common Stock issued in the Private Placement, as disclosed to the Reporting Persons by the Issuer.


SCHEDULE 13D




Comment for Type of Reporting Person:
Consists of 862,068 shares of Common Stock held of record by TCG Opportunity III LP. TCG Opportunity III GP LP is the general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to these securities. TCG Opportunity III GP LLC is the general partner of TCG Opportunity III GP LP and the ultimate general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to these securities. Peter Svennilson and Tim Kutzkey, a member of the Issuer's board of directors, are the managing members of TCG Opportunity III GP LLC and may each be deemed to share voting, investment and dispositive power with respect to these securities. Excludes warrants to purchase 431,034 shares of Common Stock held by TCG Opportunity III LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations. Based on 17,796,879 shares outstanding, comprised of (i) 11,753,558 shares of Common Stock outstanding as of August 4, 2026, as reported by the Issuer in the Form 10-Q, plus (ii) 6,043,321 shares of Common Stock issued in the Private Placement, as disclosed to the Reporting Persons by the Issuer.


SCHEDULE 13D




Comment for Type of Reporting Person:
Consists of 862,068 shares of Common Stock held of record by TCG Opportunity III LP. TCG Opportunity III GP LP is the general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to these securities. TCG Opportunity III GP LLC is the general partner of TCG Opportunity III GP LP and the ultimate general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to these securities. Peter Svennilson and Tim Kutzkey, a member of the Issuer's board of directors, are the managing members of TCG Opportunity III GP LLC and may each be deemed to share voting, investment and dispositive power with respect to these securities. Excludes warrants to purchase 431,034 shares of Common Stock held by TCG Opportunity III LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations. Based on 17,796,879 shares outstanding, comprised of (i) 11,753,558 shares of Common Stock outstanding as of August 4, 2026, as reported by the Issuer in the Form 10-Q, plus (ii) 6,043,321 shares of Common Stock issued in the Private Placement, as disclosed to the Reporting Persons by the Issuer.


SCHEDULE 13D




Comment for Type of Reporting Person:
Consists of (i) 1,386,122 shares of Common Stock held of record by TCG III LP, (ii) warrants to purchase 5,218 shares of Common Stock held by TCG III LP which are exercisable within 60 days of this Statement, (iii) 1,565,311 shares of Common Stock held of record by TCG III-A LP, (iv) warrants to purchase 5,893 shares of Common Stock held by TCG III-A LP which are exercisable within 60 days of this Statement and (v) 862,068 shares of Common Stock held of record by TCG Opportunity III LP. TCG III GP LP is the general partner of each of TCG III LP and TCG III-A LP and may be deemed to have voting, investment and dispositive power with respect to these securities. TCG Opportunity III GP LP is the general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to the securities held by TCG Opportunity III LP. TCG Opportunity III GP LLC is the general partner of TCG Opportunity III GP LP and the ultimate general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to the securities held by TCG Opportunity III LP. Peter Svennilson and Tim Kutzkey, a member of the Issuer's board of directors, are the managing partners of TCG III GP LP and the managing members of TCG Opportunity III GP LLC and may each be deemed to share voting, investment and dispositive power with respect to these securities. Excludes (a) warrants to purchase 594,625 shares of Common Stock held by TCG III LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations, (b) warrants to purchase 671,518 shares of Common Stock held by TCG III-A LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations and (c) warrants to purchase 431,034 shares of Common Stock held by TCG Opportunity III LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations. Based on 17,807,990 shares outstanding, comprised of (i) 11,753,558 shares of Common Stock outstanding as of August 4, 2026, as reported by the Issuer in the Form 10-Q, plus (ii) 6,043,321 shares of Common Stock issued in the Private Placement, as disclosed to the Reporting Persons by the Issuer, plus (iii) 5,218 shares underlying warrants to purchase shares of Common Stock held by TCG III LP which are exercisable within 60 days of this Statement, plus (iv) 5,893 shares underlying warrants to purchase shares of Common Stock held by TCG III-A LP which are exercisable within 60 days of this Statement.


SCHEDULE 13D




Comment for Type of Reporting Person:
Consists of (i) 1,386,122 shares of Common Stock held of record by TCG III LP, (ii) warrants to purchase 5,218 shares of Common Stock held by TCG III LP which are exercisable within 60 days of this Statement, (iii) 1,565,311 shares of Common Stock held of record by TCG III-A LP, (iv) warrants to purchase 5,893 shares of Common Stock held by TCG III-A LP which are exercisable within 60 days of this Statement and (v) 862,068 shares of Common Stock held of record by TCG Opportunity III LP. TCG III GP LP is the general partner of each of TCG III LP and TCG III-A LP and may be deemed to have voting, investment and dispositive power with respect to these securities. TCG Opportunity III GP LP is the general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to the securities held by TCG Opportunity III LP. TCG Opportunity III GP LLC is the general partner of TCG Opportunity III GP LP and the ultimate general partner of TCG Opportunity III LP and may be deemed to have voting, investment and dispositive power with respect to the securities held by TCG Opportunity III LP. Peter Svennilson and Tim Kutzkey, a member of the Issuer's board of directors, are the managing partners of TCG III GP LP and the managing members of TCG Opportunity III GP LLC and may each be deemed to share voting, investment and dispositive power with respect to these securities. Excludes (a) warrants to purchase 594,625 shares of Common Stock held by TCG III LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations, (b) warrants to purchase 671,518 shares of Common Stock held by TCG III-A LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations and (c) warrants to purchase 431,034 shares of Common Stock held by TCG Opportunity III LP which are not exercisable within 60 days of this Statement due to beneficial ownership limitations. Based on 17,807,990 shares outstanding, comprised of (i) 11,753,558 shares of Common Stock outstanding as of August 4, 2026, as reported by the Issuer in the Form 10-Q, plus (ii) 6,043,321 shares of Common Stock issued in the Private Placement, as disclosed to the Reporting Persons by the Issuer, plus (iii) 5,218 shares underlying warrants to purchase shares of Common Stock held by TCG III LP which are exercisable within 60 days of this Statement, plus (iv) 5,893 shares underlying warrants to purchase shares of Common Stock held by TCG III-A LP which are exercisable within 60 days of this Statement.


SCHEDULE 13D


 
The Column Group III GP, LP
 
Signature:/s/ James Evangelista
Name/Title:James Evangelista, Attorney-in-Fact
Date:10/22/2026
 
The Column Group III, LP
 
Signature:/s/ James Evangelista
Name/Title:James Evangelista, Attorney-in-Fact
Date:10/22/2026
 
The Column Group III-A, LP
 
Signature:/s/ James Evangelista
Name/Title:James Evangelista, Attorney-in-Fact
Date:10/22/2026
 
The Column Group Opportunity III, LP
 
Signature:/s/ James Evangelista
Name/Title:James Evangelista, Attorney-in-Fact
Date:10/22/2026
 
The Column Group Opportunity III GP, LP
 
Signature:/s/ James Evangelista
Name/Title:James Evangelista, Attorney-in-Fact
Date:10/22/2026
 
TCG Opportunity III GP, LLC
 
Signature:/s/ James Evangelista
Name/Title:James Evangelista, Attorney-in-Fact
Date:10/22/2026
 
Tim Kutzkey
 
Signature:/s/ James Evangelista
Name/Title:James Evangelista, Attorney-in-Fact
Date:10/22/2026
 
Peter Svennilson
 
Signature:/s/ James Evangelista
Name/Title:James Evangelista, Attorney-in-Fact
Date:10/22/2026