Total compensation cost recognized for all stock-based compensation awards in the condensed consolidated statements of operations was as follows (in thousands):


	Three Months Ended June 30,				Six Months Ended June 30,
	2026		2025		2026		2025
Research and development	$	644	$	863	$	1,187	$	1,770
General and administrative	1,549		1,801		3,108		3,873
Total stock-based compensation expense	$	2,193	$	2,664	$	4,295	$	5,643

The following is a summary of restricted stock units activity for the six months ended June 30, 2026:


	Shares	Weighted Average Grant Date Fair Value Per Share
Unvested restricted stock units as of December 31, 2025	595,730	$	9.11
Issued	—	$	—
Vested	(88,306)	$	10.22
Forfeited	(78,990)	$	8.42
Unvested restricted stock units as of June 30, 2026	428,434	$	9.11

There were no restricted stock units that contained performance-based vesting provisions granted in the six months ended June 30, 2026 and 2025. There was no expense related to the vesting of performance-based restricted stock units for the three and six months ended June 30, 2026 and 2025.

As of June 30, 2026, total unrecognized compensation expense related to unvested restricted stock units was $1.7 million, which the Company expects to recognize over a remaining weighted-average period of 1.1 years.


	Shares	Weighted Average Exercise Price		Remaining Contractual Life (years)	Aggregate Intrinsic Value
Outstanding at December 31, 2025	8,403,694	$	8.37	7.65	$	1,395
Granted	4,789,807	$	2.53
Exercised	(20,040)	$	1.80
Expired	(439,869)	$	18.75
Forfeited	(446,567)	$	2.66
Outstanding at June 30, 2026	12,287,025	$	5.94	8.27	$	9,322
Vested and expected to vest at June 30, 2026	12,287,025	$	5.94	8.27	$	9,322
Exercisable at June 30, 2026	4,510,084	$	11.62	6.67	$	1,926

As of June 30, 2026, total unrecognized compensation expense related to stock options was $14.1 million, which the Company expects to recognize over a remaining weighted-average period of 2.8 years.