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Disclosure - Summary of Accounting Pronouncements and Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 9952158 - Disclosure - Research Collaboration and License Agreements link:presentationLink link:calculationLink link:definitionLink 9952159 - Disclosure - Marketable Securities and Fair Value Measurements link:presentationLink link:calculationLink link:definitionLink 9952160 - Disclosure - Property, Equipment and Leasehold Improvements link:presentationLink link:calculationLink link:definitionLink 9952161 - Disclosure - Right-of-Use Assets and Liabilities link:presentationLink link:calculationLink link:definitionLink 9952162 - Disclosure - Accounts Payable and Accrued Liabilities link:presentationLink link:calculationLink link:definitionLink 9952163 - Disclosure - Long-Term Debt link:presentationLink link:calculationLink link:definitionLink 9952164 - Disclosure - Equity link:presentationLink link:calculationLink link:definitionLink 9952165 - Disclosure - Income Taxes link:presentationLink link:calculationLink link:definitionLink 9952166 - Disclosure - Loss Per Common Share link:presentationLink link:calculationLink link:definitionLink 9952167 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 9952168 - Disclosure - Related Party Transactions link:presentationLink link:calculationLink link:definitionLink 9952169 - Disclosure - Restructuring Activity link:presentationLink link:calculationLink link:definitionLink 9952170 - Disclosure - Segment Information link:presentationLink link:calculationLink link:definitionLink 9952171 - Disclosure - Subsequent Events link:presentationLink link:calculationLink link:definitionLink 9955511 - Disclosure - Summary of Accounting Pronouncements and Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 9955512 - Disclosure - Research Collaboration and License Agreements (Tables) link:presentationLink link:calculationLink link:definitionLink 9955513 - Disclosure - Marketable Securities and Fair Value Measurements (Tables) link:presentationLink link:calculationLink link:definitionLink 9955514 - Disclosure - Property, Equipment and Leasehold Improvements (Tables) link:presentationLink link:calculationLink link:definitionLink 9955515 - Disclosure - Right-of-Use Assets and Liabilities (Tables) link:presentationLink link:calculationLink link:definitionLink 9955516 - Disclosure - Accounts Payable and Accrued Liabilities (Tables) link:presentationLink link:calculationLink link:definitionLink 9955517 - Disclosure - Long-Term Debt (Tables) link:presentationLink link:calculationLink link:definitionLink 9955518 - Disclosure - Equity (Tables) link:presentationLink link:calculationLink link:definitionLink 9955519 - Disclosure - Loss Per Common Share (Tables) link:presentationLink link:calculationLink link:definitionLink 9955520 - Disclosure - Segment Information (Tables) link:presentationLink link:calculationLink link:definitionLink 9955521 - Disclosure - Nature of Business and Basis of Presentation (Details) link:presentationLink link:calculationLink link:definitionLink 9955522 - Disclosure - Research Collaboration and License Agreements - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955523 - Disclosure - Research Collaboration and License Agreements - Summary of Sources of Revenue (Details) link:presentationLink link:calculationLink link:definitionLink 9955524 - Disclosure - Research Collaboration and License Agreements - Schedule of Contract Balances (Details) link:presentationLink link:calculationLink link:definitionLink 9955525 - Disclosure - Research Collaboration and License Agreements - Transaction Price Allocated to Performance Obligations (Details) link:presentationLink link:calculationLink link:definitionLink 9955525 - Disclosure - Research Collaboration and License Agreements - Transaction Price Allocated to Performance Obligations (Details) link:presentationLink link:calculationLink link:definitionLink 9955526 - Disclosure - Marketable Securities and Fair Value Measurements (Details) link:presentationLink link:calculationLink link:definitionLink 9955527 - Disclosure - Property, Equipment and Leasehold Improvements - Schedule of Property, Equipment and Leasehold Improvements (Details) link:presentationLink link:calculationLink link:definitionLink 9955528 - Disclosure - Property, Equipment and Leasehold Improvements - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955529 - Disclosure - Right-of-Use Assets and Liabilities - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955530 - Disclosure - Right-of-Use Assets and Liabilities - Components of Lease Expense (Details) link:presentationLink link:calculationLink link:definitionLink 9955531 - Disclosure - Right of Use Assets and Liabilities - Schedule of Supplemental Cash Flow Information Related to Leases (Details) link:presentationLink link:calculationLink link:definitionLink 9955532 - Disclosure - Right-of-Use Assets and Liabilities - Schedule of Maturities of Operating Lease Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955532 - Disclosure - Right-of-Use Assets and Liabilities - Schedule of Maturities of Operating Lease Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955533 - Disclosure - Accounts Payable and Accrued Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955534 - Disclosure - Long-Term Debt - Schedule of Long Term Debt (Details) link:presentationLink link:calculationLink link:definitionLink 9955535 - Disclosure - Long-Term Debt - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955536 - Disclosure - Long-Term Debt - Schedule of Minimum Future Payments on Long-Term Debt (Details) link:presentationLink link:calculationLink link:definitionLink 9955537 - Disclosure - Equity - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955538 - Disclosure - Equity - Schedule of Assumptions Used to Determine Fair Value of and Stock Options Granted (Details) link:presentationLink link:calculationLink link:definitionLink 9955539 - Disclosure - Equity - Summary of Stock Option Activity (Details) link:presentationLink link:calculationLink link:definitionLink 9955540 - Disclosure - Equity - Summary of Restricted Stock (Details) link:presentationLink link:calculationLink link:definitionLink 9955541 - Disclosure - Income Taxes (Details) link:presentationLink link:calculationLink link:definitionLink 9955542 - Disclosure - Loss Per Common Share - Schedule of Basic and Diluted Loss per Common Share (Details) link:presentationLink link:calculationLink link:definitionLink 9955543 - Disclosure - Loss Per Common Share - Securities Excluded From Computation of Diluted Net Loss Per Common Share (Details) link:presentationLink link:calculationLink link:definitionLink 9955544 - Disclosure - Commitments and Contingencies (Details) link:presentationLink link:calculationLink link:definitionLink 9955545 - 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