Total compensation cost recognized for all stock-based compensation awards in the condensed consolidated statements of operations was as follows (in thousands):


	Three Months Ended March 31,
	2026		2025
Research and development	$	544	$	907
General and administrative	1,558		2,072
Total stock-based compensation expense	$	2,102	$	2,979

The following is a summary of restricted stock units activity for the three months ended March 31, 2026:


	Shares	Weighted Average Grant Date Fair Value Per Share
Unvested restricted stock units as of December 31, 2025	595,730	$	9.11
Issued	—	$	—
Vested	(34,038)	$	10.29
Forfeited	(68,003)	$	8.51
Unvested restricted stock units as of March 31, 2026	493,689	$	9.19

There were no restricted stock units that contained performance-based vesting provisions granted in the three months ended March 31, 2026. There was no expense related to the vesting of performance-based restricted stock units for the three months ended March 31, 2026 and 2025.

As of March 31, 2026, total unrecognized compensation expense related to unvested restricted stock units was $2.1 million, which the Company expects to recognize over a remaining weighted-average period of 1.34 years.

The following is a summary of stock option activity for the three months ended March 31, 2026:


	Shares	Weighted Average Exercise Price		Remaining Contractual Life (years)	Aggregate Intrinsic Value
Outstanding at December 31, 2025	8,403,694	$	8.37	7.65	$	1,395
Granted	4,424,417	$	2.52
Exercised	—	$	—
Expired	(75,582)	$	12.42
Forfeited	(127,994)	$	2.79
Outstanding at March 31, 2026	12,624,535	$	6.35	8.06	$	3,164
Vested and expected to vest at March 31, 2026	12,624,535	$	6.35	8.06	$	3,164
Exercisable at March 31, 2026	4,096,257	$	13.87	5.96	$	630

As of March 31, 2026, total unrecognized compensation expense related to stock options was $15.8 million, which the Company expects to recognize over a remaining weighted-average period of 3.00 years.