EX-FILING FEES

Calculation of Filing Fee Tables

S-3

Aura Biosciences, Inc.


Table 1: Newly Registered and Carry Forward Securities		☐ Not Applicable

Security
Type

Security
Class
Title

Fee
Calculation
or Carry
Forward
Rule

Amount
Registered

Proposed
Maximum
Offering
Price Per
Unit

Maximum
Aggregate
Offering
Price

Fee
Rate

Amount of
Registration
Fee

Carry
Forward
Form
Type

Carry
Forward
File
Number

Carry
Forward
Initial
Effective
Date

Filing Fee
Previously
Paid in
Connection
with Unsold
Securities to
be Carried
Forward

Newly Registered Securities

Equity

Common Stock, par value $0.00001 per share

457(o)

Equity

Pre-funded

warrants to purchase Common Stock

457(o)

Equity

Common stock, par value $0.00001 per share, issuable upon exercise of

Pre-Funded

Warrants

Fees to be
Paid

Unallocated (Universal) Shelf

457(o)

$49,907,450.00

0.0001381

$6,892.22

Fees
Previously
Paid

Carry Forward Securities

Total Offering Amounts:

$49,907,450.00

$6,892.22

Total Fees Previously Paid:

$ 0.00

Total Fee Offsets:

$ 0.00

Net Fee Due:

$6,892.22

Offering Note

The amount to be registered consists of up to $49,907,450.00 of an indeterminate amount of common stock, par

val

ue $0.00001 per share (the “Common Stock”),

pre-funded

warrants to purchase shares of the Company’s stock

(“Pre-Funded

Warrants”) and shares issuable upon the exercise of

Pre-Funded

Warrants. Any securities registered hereunder may be sold separately or together with other securities registered hereunder. Pursuant to Rule 416 under the Securities Act of 1933, as amended, this registration statement also covers any additional securities that may be offered or issued in connection with any stock split, stock dividend or pursuant to anti-dilution provisions of any of the securities. Separate consideration may or may not be received for securities that are issuable upon conversion, exercise or exchange of other securities.

The proposed maximum offering price per security will be determined from time to time by the registrant in connection with the issuance by the registrant of the securities registered hereunder and is not specified as to each class of security pursuant to General Instruction II.D. of Form

S-3

under the Securities Act of 1933, as amended.

Includes such indeterminate amount of common stock as may be issued from time to time upon exercise of

Pre-Funded

Warrants registered hereby, as the case may be.

Pre-Funded

Warrants may be sold separately or together with any of the securities registered hereby and may be exercisable for shares of common stock registered hereby. Because the

Pre-Funded

Warrants will provide a right only to purchase such securities offered hereunder, no additional registration fee is required.

Table 2: Fee Offset Claims and Sources

☑

Not Applicable

Registrant
or Filer
Name

Form
or
Filing
Type

File
Number

Initial
Filing
Date

Filing
Date

Fee
Offset
Claimed

Security
Type
Associated
with Fee
Offset
Claimed

Security
Title
Associated
with Fee
Offset
Claimed

Unsold
Securities
Associated
with Fee
Offset
Claimed

Unsold
Aggregate
Offering
Amount
Associated
with Fee
Offset
Claimed

Fee
Paid
with
Fee
Offset
Source

Rules 457(b) and

0-11(a)(2)

Fee Offset Claims

N/A

Fee Offset Sources

N/A

Rule 457(p)

Fee Offset Claims

N/A

Fee Offset Sources

N/A

Table 3: Combined Prospectuses

☑

Not Applicable

Security
Type

Security
Class Title

Amount of Securities
Previously Registered

Maximum Aggregate
Offering Price of
Securities Previously
Registered

Form Type

File
Number

Initial
Effective
Date

N/A