Exhibit 99.1

EXHIBIT A
JOINT FILING AGREEMENT
The undersigned hereby agree that the statement on Schedule 13G with respect to the Common Stock of Keros Therapeutics, Inc. dated as of May 14, 2025 is, and any further amendments thereto signed by each of the undersigned shall be, filed on behalf of each of the undersigned pursuant to and in accordance with the provisions of Rule 13d-1(k)(2) under the Securities Exchange Act of 1934, as amended.
Dated: May 14, 2025
MADISON AVENUE INTERNATIONAL LP


By: /s/ Eli Samaha
    Eli Samaha, as Manager of Madison Avenue GP, LLC, the General Partner of Madison Avenue International LP


MADISON AVENUE PARTNERS, LP


By: /s/ Eli Samaha
    Eli Samaha, as Managing Member of EMAI Management, LLC, the General Partner of Madison Avenue Partners, LP


EMAI MANAGEMENT, LLC


By: /s/ Eli Samaha
    Eli Samaha, as Managing Member


MADISON AVENUE GP, LLC


By: /s/ Eli Samaha
    Eli Samaha, as Manager


CARAWAY JACKSON INVESTMENTS, LLC


By: /s/ Eli Samaha
    Eli Samaha, as Member


/s/ Eli Samaha
Eli Samaha